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(Click to enlarge)

Just a note that this seems to be occurring again with custom zones. 10.0.30 fresh install on live server running php 7.4. If this isn't a bug, then perhaps a tutorial on breadcrumbs for custom zones running catalogues transparently from site/ would be useful because the closest I can get is for the Zone to be shown in the breadcrumbs on catalogue entries, I cannot get the Zone in the breadcrumbs on the category page. On the entry page, we lose the parent page in the breadcrumbs as with this previous issue.

I set up a transparent redirect so catalogues could run out of the collaboration zone then added this ABOVE the default catalogue breadcrumbs.xml rules and it worked ok.

<substitution match_key="collaboration:catalogues:index:tree=0" final="true"> <!-- Skip catalogues list. -->
<link label="{!HOME}">:</link>
<link label="Collaboration">collaboration</link>
</substitution>
<substitution match_key="collaboration:catalogues:category"> <!-- Skip catalogues list. This rule won't run for non-tree catalogues as the above rule terminates first; i.e. it is only for tree catalogues -->
<link label="{!HOME}">:</link>
<link label="Collaboration">collaboration</link>
</substitution>
<substitution match_key="collaboration:catalogues:atoz"> <!-- Skip catalogues list. -->
<link label="{!HOME}">:</link>
<link label="Collaboration">collaboration</link>
</substitution>

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Above might be the trick I'm missing, I added them below the currently defined breadcrumbs.

Still not happening for me with a custom zone, collaboration isn't a brand new zone. I can get them working on pages, just not catalogues running in a custom zone.

Works for me with a custom zone too. Catch me on IRC to discuss.